This is after finding out the other day what these buttons did. I've been working on a vtuber model, and was looking for a tool that would mirror my brush strokes horizontally. Turns out, it's been sitting up there for the past 10 years and I just never paid any attention to it. Go figure.
I should probably make clear, these buttons mirror your brush strokes across a line. Whatever you draw on one side will also be drawn on the other. The shortcut I found tonight flips the canvas. Two different mirror tools.